Peer-reviewed journals


Gallou A, Jump AS, Lynn JS, Field R, Irl SDH, Steinbauer MJ, ... & Grytnes JA (2023). Diurnal temperature range as a key predictor of plants’ elevation ranges globally. Nature Communications, 14, 7890.

Hanz DM, Cutts V, Barajas‐Barbosa MP, Algar A, Beierkuhnlein C, Collart F, ... & Irl SDH (2023). Effects of climate change on the distribution of plant species and plant functional strategies on the Canary Islands. Diversity and Distributions, 29, 1157-1171.

Cutts V, Hanz DM, Barajas‐Barbosa MP, Schrodt F, Steinbauer MJ, Beierkuhnlein C, ...Irl SDH... & Algar, A. C. (2023). Links to rare climates do not translate into distinct traits for island endemics. Ecology Letters, 26, 504-515.

Kienle DR, Irl SDH, & Beierkuhnlein C (2023). Mass elevation effect and continentality have a stronger impact on global treelines than spatial isolation. Global Ecology and Biogeography, 32, 1087-1097.

Walentowitz A, Ferreira‐Arruda T, Irl SDH, Kreft H, & Beierkuhnlein C (2023). Disentangling natural and anthropogenic drivers of native and non‐native plant diversity on North Sea islands. Journal of Biogeography.


Eibes PM, Schaffrath F, Oldeland J, Thormählen W, Schmiedel U, Irl SDH (2022) Testing the concept of edaphism for the quartz island flora of the Knersvlakte, South Africa. South African Journal of Botany 151, 555-564.

Hanz DM, Cutts V, Barajas‐Barbosa MP, Algar AC, Beierkuhnlein C, Fernández‐Palacios JM, Field R, Kreft H, Steinbauer MJ, Weigelt P, Irl SDH (2022) Climatic and biogeographical drivers of functional diversity in the flora of the Canary Islands. Global Ecology and Biogeography 31. 1313-1331.

Oldeland J, Eibes PM, Irl SDH, Schmiedel U (2022) Do image resolution and classifier choice impact island biogeographical parameters of terrestrial islands? Transactions in GIS 26, 2004-2022.

Kienle D, Walentowitz A, Sungur L, Chiarucci A, Irl SDH, Jentsch A, Ole R Vetaas, Field R, Beierkuhnlein C (2022) Geodiversity and biodiversity on a volcanic island: the role of scattered phonolites for plant diversity and performance. Biogeosciences 19, 1691-1703.

Hanz DM, Beloiu M, Wipfler R, Beierkuhnlein C, Field R, Jentsch A, Vetaas OR, Irl SDH (2022) High species turnover and low intraspecific trait variation in endemic and non‐endemic plant species assemblages on an oceanic island. Journal of Vegetation Science 33, e13120.


Cervellini M, Di Musciano M, Zannini P, Fattorini S, Jiménez‐Alfaro B, Agrillo E, Attorre F, Angelini P, Beierkuhnlein C, Casella L, Field R, Fischer JC, Genovesi P, Hoffmann S, Irl SDH, Nascimbene J, Rocchini D,  Steinbauer MJ, Vetaas OR, Chiarucci A (2021) Diversity of European habitat types is correlated with geography more than climate and human pressure. Ecology and Evolution 24, 18111-18124.

Weiser F, Sauer A, Gettueva D, Field R, Irl SDH, Vetaas O, Chiarucci A, Hoffmann S, Fermández-Palacios JM, Otto R, Jentsch A, Provenzale A, Beierkuhnlein C (2021) Impacts of forest fire on understory species diversity in the Canary pine ecosystems on the island of La Palma. Forests 12, 1638.

Fernández-Palacios JM, Kreft H, Irl SDH, Norder S, Ah-Peng C, Borges PA, Burns KC, de Nascimento L, Meyer JY, Montes E & Drake DR (2021). Scientists’ Warning-The Outstanding Biodiversity of Islands is in Peril. Global Ecology and Conservation, e01847.


Eibes PM, Oldeland J, Irl SDH, Twerski A, Kühne N, Schmiedel U (2021). Partitioned beta diversity patterns of plants across sharp and distinct boundaries of quartz habitat islands. Journal of Vegetation Science, 32:e13036, DOI:

Eibes PM, Eisenbacher J, Beierkuhnlein C, Chiarucci A, Field R, Jentsch A, Köhler T, Veetas OR, Irl SDH (2021). Co-occurrence frequency in vegetation patches decreases towards the harsh edge along an arid volcanic elevational gradient. Frontiers of Biogeography, 13.3, e49743, DOI:

Cutts V, Hanz DM, Barajas-Barbosa MP, Algar AC, Steinbauer MJ, Irl SDH, Kreft H, Weigelt P, Fernández-Palacios JM, Field R (2021) Scientific floras can be reliable sources for some trait data in a system with poor coverage in global trait databases. Journal of Vegetation Science.

Irl SDH, Schweiger AH, Steinbauer MJ, Ah-Peng C, Arévalo JR, Beierkuhnlein C, Chiarucci A, Daehler CC, Fernández-Palacios JM, Flores O, Kueffer C, Madera P, Otto R, Schweiger JMI, Strasberg D, Jentsch A (2021) Human impact, climate and dispersal strategies determine plant invasion on islands. Journal of Biogeography.



Lenzner B, Latombe G, Capinha C, Bellard C, Courchamp F, Diagne C, Dullinger S, Golivets M, Irl SDH, Kühn I, Leung B, Liu C, Moser D, Roura-Pascual N, Seebens H, Turbelin A, Weigelt P & Essl F (2020) What will the future bring for biological invasions on islands? An expert-based assessment. Frontiers in Ecology and Evolution, 8, 280. DOI:

Flantua S, Payne D, Borregaard M, Beierkuhnlein C, Steinbauer MJ, Dullinger S, Essl F, Irl SDH, Kienle D, Kreft H, Lenzner B, Norder S, Rijsdijk K, Rumpf S, Weigelt P, Field R (2020) Snapshot isolation and isolation history challenge the analogy between mountains and islands used to understand endemism. Global Ecology and Biogeography, DOI:

Irl SDH, Obermeier A, Beierkuhnlein C, Steinbauer MJ (2020) Climate controls plant life form patterns on a high-elevation oceanic island. Journal of Biogeography, DOI:

Cervellini M, Zannini P, Di Musciano M, Fattorini S, Jiménez-Alfaro B, Rocchini D, ... Irl SDH,... & Chiarucci A (2020) A grid-based map for the Biogeographical Regions of Europe. Biodiversity Data Journal, 8, e53720.

Schweiger AH, Irl SDH, Svenning JC, Higgins SI (2020) Dynamic management needs for long‐lived, sporadically recruiting plant species in human‐dominated landscapes. Plants, People, Planet, 2 , 186-200.

Cutts V, Katal N, Löwer C, Algar AC, Steinbauer MJ, Irl SDH, Beierkuhnlein C, Field R (2020) The effect of small-scale topography on patterns of endemism within islands. Frontiers of Biogeography, 11, e43737.


Hoffmann S, Irl SDH, Beierkuhnlein C (2019) Predicted climate shifts within terrestrial protected areas worldwide. Nature Communications, 10, 4787.

Walentowitz A, Irl SDH, Acevedo Rodríguez AJ, Palomares-Martínez Á, Vetter V, Zennaro B, MEdina FM, Beierkuhnlein C (2019) Graminoid Invasion in an Insular Endemism Hotspot and its Protected Areas. Diversity, 11, 192.

Irl SDH, Schweiger AH, Hoffmann S, Beierkuhnlein C, Hartmann H, Pickel T, Jentsch A (2019). Spatiotemporal dynamics of plant diversity and endemism during primary succession on an oceanic‐volcanic island. Journal of Vegetation Science, 30, 587-598,

Bernd Josef Berauer, Peter A. Wilfahrt, Mohammed A. S. Arfin-Khan, Pia M. Eibes, Andreas Von Heßberg, Johannes Ingrisch, Michael Schloter, Max A. Schuchardt & Anke Jentsch(2019). Low resistance of montane and alpine grasslands to abrupt changes in temperature and precipitation regimes. Arctic, Antarctic, and Alpine Research,51:1,215-231,DOI: 10.1080/15230430.2019.1618116

Cubas, J.*, Irl, S.D.H.*, Villafuerte, R., Bello-Rodríguez, V., Rodríguez-Luengo, J.L., del Arco, M., Martín-Esquivel, J.L., & González-Mancebo, J.M. (2019). Endemic plant species are more palatable to introduced herbivores than non-endemics. Proceedings of the Royal Society: B, 286, 20190136.

* Equal contribution


Weigel, R., Irl, S.D.H., Treydte, K., Beierkuhnlein, C., Berels, J., Field, R., Miranda, J., Steinbauer, A., Steinbauer, M.J., Jentsch, A. (2018). A novel dendroecological method finds a non-linear relationship between elevational and seasonal growth continuity on an island with trade wind-induced water availability. AoB PLANTS, 10, ply070; doi: 10.1093/aobpla/ply070

Hanz, D.M., Böhning-Gaese, K., Ferger, S.W., Fritz, S.A., Neuschulz, E.L., Quitián, M., Santillán, V., Töpfer. T. & Schleuning M. (2018). Functional and phylogenetic diversity of bird assemblages are filtered by different biotic factors on tropical mountains. Journal of Biogeography. doi:10.1111/jbi.13489
Cubas, J; Martín-Esquivel, JL; Nogales, M; Irl, SDH; Hernández Hernández, R; López-Darias, M; Marrero-Gómez, M; Del Arco, M; González-Mancebo, JM (2018): Contrasting effects of invasive rabbits on endemic plants driving vegetation change in a subtropical alpine insular environment, Biological Invasions, 20, 793–807: doi:10.1007/s10530-017-1576-0
Hoffmann, S; Schmitt, T; Chiarucci, A; Irl, SDH; Rocchini, D; Vetaas, OR; Tanase, MA; Mermoz, S; Bouvet, A; Beierkuhnlein, C (2018): Remote sensing of beta diversity: evidence from plant communities in a semi‐natural system, Applied Vegetation Science: doi:10.1111/avsc.12403


 Irl, SDH*; Schweiger, A*; Medina, F M; Fernández-Palacios, JM; Harter, D; Jentsch, A; Provenzale, A; Steinbauer, MJ; Beierkuhnlein, C (2017): An island view of endemic rarity – environmental drivers and consequences for nature conservation, Diversity and Distributions, 23, 1132-1142. doi: 10.1111/ddi.12605
* Equal contribution
Steinbauer, MJ*, Irl, SDH*, González-Mancebo, JM, Breiner, F, Hernández, RH, Hopfenmüller, S, Kidane Ogubamichael, Y, Jentsch, A, Beierkuhnlein, C (2017): Plant invasion and speciation along elevational gradients on the oceanic island La Palma, Canary Islands., Ecology and Evolution, 7, 771–779. doi:10.1002/ece3.2640
* Equal contribution


Irl, SDH (2016): Plant diversity on high elevation islands – drivers of species richness and endemism, Frontiers of Biogeography, 8, e29717: doi:10.21425/F58329717
Irl, SDH; Anthelme, F; Harter, D; Jentsch, A; Lotter, E; Steinbauer, M; Beierkuhnlein, C (2016): Patterns of island treeline elevation – a global perspective, Ecography, 39, 427-436: doi:10.1111/ecog.01266
Schweiger, A; Irl, SDH; Steinbauer, M; Dengler, J; Beierkuhnlein, C (2016): Optimizing sampling approaches along ecological gradients, Methods in Ecology and Evolution, 7, 463-471: doi:10.1111/2041-210X.12495
Steinbauer, M; Beierkuhnlein, C; Arfin Khan, MAS; Harter, D; Irl, SDH; Jentsch, A; Schweiger, A; Svenning, JC; Dengler, J (2016): How to differentiate facilitation and environmentally driven coexistence, Journal of Vegetation Science, 27, 1071-1079: doi:10.1111/jvs.12441
Steinbauer, M; Field, R; Fernández-Palacios, JM; Irl, SDH; Otto, Rüdiger; Schaefer, H; Beierkuhnlein, C (2016): Biogeographic ranges do not support niche theory in radiating Canary Island plant clades, Global Ecology and Biogeography, 25(7), 792-804: doi:10.1111/geb.12425
Steinbauer, M; Field, R; Grytnes, JA; Trigas, P; Ah-Peng, C; Attorre, F; Birks, HJB; Borges, PAV; Cardoso, P; Chou, C-H; De Sanctis, M; Sequera, MM; Duarte, MC; Elias, RB; Fernandez-Palacios, JM; Gabriel, R; Gereau, RE; Gillespie, RG; Greimler, J; Harter, D; Huang, T-J; Irl, SDH; Jeanmonod, D; Jentsch, A; Jump, AS; Kueffer, C; Nogué, S; Otto, R; Price, J; Romeiras, MM; Strasberg, D; Stuessy, T; Svenning, JC; Vetaas, OR; Beierkuhnlein, C (2016): Topography-driven isolation, speciation and a global increase of endemism with elevation, Global Ecology and Biogeography, 25(9), 1097–1107: doi:10.1111/geb.12469


Harter, D; Irl, SDH; Seo, B; Steinbauer, M; Gillespie, RG; Triantis, KA; Fernandez-Palacios, JM; Beierkuhnlein, C (2015): Impacts of global climate change on the floras of oceanic islands - Projections, implications and current knowledge, Perspectives in Plant Ecology, Evolution and Systematics, 17(2), 160–183: doi:10.1016/j.ppees.2015.01.003
Irl, SDH; Harter, D; Steinbauer, M; Puyol Gallego, D; Fernández-Palacios, JM; Jentsch, A; Beierkuhnlein, C (2015): Climate vs. topography – spatial patterns of plant species diversity and endemism on a high-elevation island, Journal of Ecology, 103, 1621-1633: doi:10.1111/1365-2745.12463


Irl, SDH; Steinbauer, M; Epperlein, L; Harter, D; Jentsch, A; Pätz, S; Wohlfahrt, C; Beierkuhnlein, C (2014): The hitchhiker's guide to island endemism - biodiversity and endemic perennial plant species in roadside and surrounding vegetation, Biodiversity and Conservation, 23, 2273–2287: doi:10.1007/s10531-014-0722-6
Irl, SDH; Steinbauer, M; Messinger, J; Blume-Werry, G; Palomares-Martínez, A; Beierkuhnlein, C; Jentsch, A (2014): Burned and devoured - Introduced herbivores, fire and the endemic flora of the high elevation ecosystem on La Palma, Canary Islands, Arctic Antarctic and Alpine Research, 46(4), 859-869: doi:10.1657/1938-4246-46.4.859
Warren, SD; Alt, M; Olsona, KD; Irl, SDH; Steinbauer, M; Jentsch, A (2014): The relationship between the spectral diversity of satellite imagery, habitat heterogeneity, and plant species richness, Ecological Informatics, 24, 160-168: doi:0.1016/j.ecoinf.2014.08.006


Irl, SDH; Jentsch, A; Walther, GR (2013): Papaver croceum LEDEB. - a rare example of an alien species in alpine environments of the Upper Engadine, Switzerland, Alpine Botany, 123(1), 21-30: doi:10.1007/s00035-013-0111-x

Steinbauer, M; Irl, SDH; Beierkuhnlein, C (2013): Elevation-driven ecological isolation promotes diversification on Mediterranean islands, Acta Oecologica, 47, 52-56: doi:10.1016/j.actao.2012.11.004


Irl, SDH; Steinbauer, M; Babel, W; Beierkuhnlein, C; Blume-Werry, G; Messinger, J; Palomares-Martínez, A; Strohmeier, S; Jentsch, A (2012): An 11-yr exclosure experiment in a high-elevation island ecosystem: Introduced herbivore impact on shrub species richness, seedling recruitment and population dynamics, Journal of Vegetation Science, 23(6), 1114-1125: doi:10.1111/j.1654-1103.2012.01425.x

Leutner, B; Steinbauer, M; Müller, CM; Früh, AJ; Irl, SDH; Jentsch, A; Beierkuhnlein, C (2012): Mosses Like It Rough - Growth Form Specific Responses of Mosses, Herbaceous and Woody Plants to Micro-Relief Heterogeneity, Diversity, 4, 59-73: doi:10.3390/d4010059

Non peer-reviewed journals and series

Beierkuhnlein, C; Irl, SDH (2017): Schatzinseln in Gefahr? Klimawandel und Naturschutz auf ozeanischen Inseln, Spektrum - Universität Bayreuth, 13, 46-49 - pdf link

Irl, SDH; Beierkuhnlein, C (2011): Distribution of endemic plant species on an oceanic island – a geospatial analysis of La Palma (Canary Islands), Procedia Environmental Sciences(7), 170-175

Books and book chapters

Irl SDH & Schweiger AH (2020) Hochgebirge: Hotspots der Biodiversität. In: Lozán J. L., S.-W. Breckle, H. Grassl et al. (Hrsg.). Warnsignal Klima: Hochgebirge im Wandel. S. 62-69. Online: doi:10.2312/warnsignal.klima.hochgebirge-im-wandel.09.

Geller-Grimm, F., Lerp, H., Hanz, D.M., Manz, C., Fickert, B., Kaiser, U., Lotz-Winter, H., Piepenbring, M. (2017): Pilze: Nahrung, Gift und Mythen / Fungi: Food, Poison and Mythology, Museum Wiesbaden. - pdf link

Steinbauer, M; Schweiger, A; Irl, SDH (2016): Patterns in Biogeography, Encyclopedia of Evolutionary Biology. vol. 1, Oxford: Academic Press, 221–230:

Irl, SDH (2014): Patterns and Disturbance-Induced Drivers of Plant Diversity and Endemism on High Elevation Islands, Dissertation - pdf link